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echo: evolution
to: All
from: Tim Tyler
date: 2004-07-09 14:34:00
subject: Re: Absolute or just rela

Name And Address Supplied 
wrote or quoted:
> Tim Tyler  wrote in message
news:...
> > Name And Address Supplied
 wrote or quoted:
> > > "John Edser"  wrote in
message news:...

> > > > JE:-
> > > > The biggest problem with the concept of "1" 
> > > > as relative fitness comparison maximum is that
> > > 
> > > 1 is not the maximum; the average is 1.
> > 
> > Relative fitness is often defined as being a value between 0 and 1 - e.g.:
> > 
> > ``Relative Fitness
> > 
> >   The selection coefficient is used to model the health of an individual 
> >   relative to the health of the rest of the population. 
> 
> This is sensible
> 
> >   The relative fitness of an individual, denoted w, is defined to be 
> >   w=(1-s)^n where n is the number of mutations the individual carries. 
> >   Note that w is also a value between 0 and 1, where 1 indicates a 
> >   perfectly healthy individual (no mutations) and values less than 1 are 
> >   relatively unhealthy individuals.''
> 
> This contradicts the previous statement. (1-s)^n is not the fitness of
> the class of individuals with n mutations relative to "the rest of the
> population", but rather it is relative to the zero class. If we really
> were to measure the fitness relative to the population as a whole,
> then some of the population members would have relative fitness
> exceeding unity.

The figures given on the page appear to be either expected or average
fitnesses (for individuals with that genome) - relative to the
success (or average success) of an unmutated individual.

If mututions are all deleterious (which is a good first approximation)
then such fitnesses are likely to lie between 0 and 1.

Anyway - thanks for making it clear that your relative fitness -
and the relative fitness of the example above - are indeed
relative to different objects.
-- 
__________
 |im |yler  http://timtyler.org/  tim{at}tt1lock.org  Remove lock to reply.
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